Giant Otter Bibliography

 Assembled by Nicole Duplaix, Jessica Groenendijk, Christof Schenck, Elke Staib and Sheila Sykes-Gatz

NOTE: This list of references is not meant to be exhaustive or complete but to be a working tool for persons who are interested in Giant otters but who do not have access to a large university library. We welcome any additions or corrections: Nicole.Duplaix@oregonstate.edu

Early Descriptions, Explorers’ Reports (1777-1952)

 Allen, J. A. (1910). Mammals from the Caura district of Venezuela. Bull. Am. Mus. Nat. Hist., 28: 146.

André, E., (1904). A Naturalist in the Guianas. London: Smith, Elder & Co.

Azara, F. de (1802) Apuntamientos para la Historia Natural des los Quadrupedos de Paraguay. 304-309. Madrid. La Viuda de Ibarra.

Beebe, W., G.I. Hartley & P.G. Howes (1917) Tropical Wildlife in British Guiana. New York: N. Y. Zoological Society.

Brown, C.B. (1876) Canoe and Camp Life in Bristish Guiana. London: E. Stanford.

Desmarest, A. G. (1817) Nouveau Dictionnaire d’Histoire Naturelle. 18: 208-219

Desmarest, A. G. (1820) Mammalogie. 1. Paris

Gmelin, CJ. F. ed. (1788) Linné, C. Systema Naturae 1: 93 (13th ed.). Brussels: Lemaire.

Gray, J.E. (1868) Observations on the Margin-tailed Otter Pteronura sandbachii. Proc. Zool. Soc. London 1868; 60-66.

Gray, J.E. (1869) Catalogue of Carnivorous Mammalia.: 80-81, 100-119. London: British Museum.

Nehring, A. (1887) Uber die Gray’schen Fischotter-gattungen Lutronectes, Lontra und Pteronura. Sber. Ges. Naturf. Freude Berlin. 1899:21-25

Nehring, A. (1901) Einige Notizen ueber die Lutra (Pteronura) paranensis des hiesigen Zoologischen Gartens. Sber. Ges. Naturf. Freude Berlin, 1901: 133-135.

Pohle, H. (1920) Die Unterfamilie der Lutrinae. Arch. Naturgesch, 85 Ab. H. Helft 9: 1-247.

Quelch, J. J. (1910) Animal Life in British Guiana. Georgetown: Argosy.

Rengger, J.R. (1830) Naturgeschichte der Säugetiere von Paraguay: 128-138. Basel: Schweighausersche.

Sanderson, I.T. (1950) A brief review pf the mammals of Suriname, based upon a collection made in 1938. Proc. Zool. Soc. London, 119: 755-789.

Vieira, C. O. da C. (1952) Notas sobre os mamiferos obtidos pela expedicao do Instituto Butanta as Rio das Mortes e Serra do Roncador. Papeis Dept. Zool. S. Paulo 10:117.

Zimmermann, E. A. G. (1777) Specimen Zoologiae Geographicae, Quadrupedum: 303-304, 485. Leyden.

Field Studies and other reports, by country

Argentina

Chehebar, C. (1991). Searching for the giant otter in northeastern Argentina. IUCN Otter Spec. Group Bull. 6/1991; pp. 17-18

Beccaceci, M.D. & Waller, T. (2000). Presence of the Giant Otter, Pteronura brasiliensis, in the Corrientes Province, Argentina. IUCN Otter Spec. Group Bull. 17(1)2000; pp. 31-33

Parera, A.F. (1992). Present Knowledge on the Giant Otter in Argentina. IUCN Otter Spec. Group Bull. 7/1992; pp. 19-21

Bolivia

Anderson, S. (1997). Mammals of Bolivia, Taxonomy and Distribution. Bull. Am. Mus. Nat. Hist., No. 231; pp. 343 – 344

Dunstone, N. (1987). Quest for the River Wolf - Report 1987. Durham University Expedition to Bolivia; 77 pp.

Ergueta, P. & Morales, C. de [eds.] (1996). Libro rojo de los vertebrados de Bolivia. CDC - Bolivia; pp. 171-172

Gonzales, E.R. (1997). Ecoetología de la londra (Pteronura brasiliensis), en la Reserva de Producción del Bajo Paraguá. Tesis de Grado, Universidad Autonoma “Gabriel René Moreno”, Santa Cruz de la Sierra, Bolivia; 118 pp.

Van Damme, P.A., S. Ten, R. Wallace, L. Painter, A. Taber, R. Gonzáles Jiménes, A. Fraser, D. Rumiz, C. Tapia, H. Michels, Y. Delaunoy, J. Saravia, J. Vargas, L. Torres. 2002. Distribución y estado de las poblaciones de londra Pteronura brasiliensis en Bolivia. Revista Boliviana de Ecologia y Conservación Ambiental, 12:111-134.

Van Damme, P.A., Wallace, R., Swaenepoel, K., Painter, L., Ten, S., Taber, A., Gonzales Jimenes, R., Saravia, I. Fraser, A., Vargas, J. 2002. Distribution and population status of the giant otter Pteronura brasiliensis in Bolivia. IUCN Otter Spec. Group Bull., 19 (2): 87-96.

Van Damme, P., R.B. Wallace. 2005. Considerations on measuring giant otter (Pteronura brasiliensis) relative abundance for conservation planning. Revista Boliviana de Ecologia y Conservación Ambiental, 17:65-76. PDF 633KB

Brazil

Carter, S.K. & Rosas, F.C.W. (1997). Biology and conservation of the Giant Otter Pteronura brasiliensis. Mammal Rev. 1997, Vol. 27, No.1, pp. 1-26 PDF 6MB

Evangelista, E., (2006). Censimento e Osservazioni Comportamentali sulla Lontra Gigante (Pteronura brasiliensis) nella Riserva Xixuau-Xipariná, Roraima, Brasile. Tesi di Laurea, Università Di Roma “La Sapienza”, Roma, Italia, pp. 113

Lima, D., Marmontel, M., Calvimontes, J., Mattos, G.E. & Rosas, F. C. W. (2005) . Acompanhamento de grupos remanescentes de ariranha (Pteronura brasiliensis) na Reserva de Desenvolvimento Sustentável Amanã, Amazonas.. In: III Congresso Brasileiro de Mastozoologia, 2005, Aracruz. Livro de Resumos Congresso Brasileiro de Mastozoologia, 2005.  

Machado, M. R. B. & Rosas, F. C. W. (2003). Vocal Repertoire in Pteronura brasiliensis (Mammalia: Carnivora): Preliminary analysis. In: XV Biennial Conference on the Biology of Marine Mammals, 2003, Greensboro, USA. Abstracts of the XV Biennial Conference on the Biology of Marine Mammals, 2003.

Mattos, G. E., Rosas, F. C. W., Lazzarini, S. M. & Picanço, M. C. L. (2004). Caracterização de tocas e seu uso por ariranhas (Pteronura brasiliensis) no lago da UHE Balbina, Amazonas, Brasil. In: XI Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2004, Quito. Livro de Resumos da XI Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2004. p. 135-135. PDF 405KB

Mattos, G. E., Rosas, F. C. W., Picanço, M. C. L.&; Lazzarini, S. M. (2002). Utilização do lago da Hidrelétrica de Balbina (Amazonas, Brasil) por Pteronura brasiliensis. In: X Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2002, Valdivia. Resumos X Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2002. p. 74-75. PDF 481KB

Ribas Pereira, C. (2004). Desenvolvimento de um Programa de Monitoramento em Longo Prazo das Ariranhas (Pteronura brasiliensis) no Pantanal Brasileiro (Long-term monitoring study of Giant otters in the Pantanal, Brazil). MSc dissertation, 68 pages PDF 1.6MB

Rosas, F. C. W. Técnicas de campo que podem auxiliar no estudo de populações naturais de ariranhas (Pteronura brasiliensis). Boletim Informativo da Sociedade Brasileira de Mastozoologia, SBMz Porto Alegre, p. 04 - 04. PDF 1MB

Rosas, F. C. W. & Mattos, G. E. (2003) Notes on giant otter (Pteronura brasiliensis) behavior in the lake of the Balbina hydroelectric power station, Amazonas, Brazil. The Latin American journal of aquatic mammals, v. 2, n. 2, p. 127-129, 2003. PDF 804KB

Rosas, F. C. W. & Mattos, G. E. (2003). Natural deaths of giant otters (Pteronura brasiliensis) in Balbina Hydroelectric lake, Amazonas, Brazil. IUCN Otter Specialist Group bulletin, Amsterdam, v. 20, n. 2, p. 62-64, 2003. PDF 633KB

Rosas, F. C. W., Mattos, G. E. & Cabral, M. M. M. (2006). Densidade de tocas de ariranhas (Pteronura brasiliensis) em um trecho do Rio Pitinga no Reservatório da UHE Balbina, AM, Brasil. In: I Congresso Sulamericano de Mastozoologia, 2006, Gramado. Livro de Resumos. Porto Alegre : UFRGS Gráfica, 2006. PDF 788KB

Rosas, F. C. W., Matos, G. E. & Cabral, M. M. M. (2005) Giant otter (Pteronura brasiliensis) in an Amazonian hydroelectric lake. In: XVI Biennial Conference on the Biology of Marine Mammals, 2005, San Diego, CA - USA. Livro de resumos da XVI Biennial Conference on the Biology of Marine Mammals, 2005.

Rosas, F. C. W., Mattos, G. E., Lazzarini, S. M. & Picanco, M. C. L. (2004) . Sazonalidade reprodutiva de ariranha (Pteronura brasiliensis) no lago da Usina Hidrelétrica de Balbina, Amazonas, Brasil. In: XI Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2004, Quito. Livro de Resumos da XI Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2004. p. 94-94. PDF 396KB

Schweizer, J. (1986). Ocorrencia e alguns dados biologicos de ariranha (Pteronura brasiliensis) no Rio Negro. In Reuniao de trabalho de especialistas en mamiferos aquaticos da America do Sul. 20. Resumos. p. 48.

Zuanon, J. A., Mattos, G. E. & Rosas, F. C. W. (2002). Hábitos alimentares de Pteronura brasiliensis no lago da Hidrelétrtica de Balbina, Amazônia Central, Brasil. In: X Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2002, Valdivia. Resumos X Reunión de Trabajo de Especialistas en Mamíferos Acuáticos de América del Sur, 2002. p. 119. PDF 528KB

Colombia

Botello, J.C. (2000). Ecología y comportamiento del lobo de río Pteronura brasiliensis en la región del bajo río Apaporis, Amazonía colombiana. Tesis de grado. Universidad del Valle. Cali – Colombia. 64 p.

Defler, T.R. ( ). A Census of Pteronura brasiliensis in El Tuparro National Park and Environs, Colombia. Presented to National Parks System of Colombia INDERENA; 20 pp.

Defler, T.R. (1986). The giant river otter in El Tuparro National Park, Colombia. Oryx Vol 20 No 2/April; pp. 87-88

Donadio, A. (1978). Some Comments on Otter Trade and Legislation in Colombia. In: Proceedings First Working Meeting of the IUCN Otter Specialist Group, Paramaribo, Suriname, March 1977, N. Duplaix (ed.); pp. 34-42

Gómez, J.R. & Jorgenson, J. (1999). An Overview to the Giant Otter – Fisherman Problem in the Orinoco Basin of Colombia. IUCN Otter Spec. Group Bull. 16 (2)1999: 90-96

Gómez, J.R. (1999). Ecología alimentaria de la Nutria Gigante (Pteronura brasiliensis) en el bajo Río Bita (Vichada, Colombia). Tesis, Universidad Javeriana, Santafé de Bogotá, Colombia.

Trujillo, F. & J.C. Botello. (2002). Pteronura brasiliensis (Omelin 1788). En: Libro rojo de los mamíferos de Colombia. Serie Libros Rojos de Especies Amenazadas de Colombia. Conservación Internacional Colombia, Instituto de Ciencias Naturales – Universidad Nacional de Colombia, Ministerio del Medio Ambiente. Bogotá, Colombia, 2005.

Velasco Gómez, D.M. (2004). Valoración biológica y cultural de la nutria gigante en el área de influencia de Puerto Carreño, Vichada, Colombia (ríos Orinoco, Bita y caños Juriepe y Negro). Thesis, Carrera De Ecología, Pontificia Universidad Javeriana Facultad De Estudios Ambientales Y Rurales Bogota, Colombia. PDF 2MB

Velasco, D.M. (2005). Investigación en Guainia: Estudio preliminar sobre el estado de conservación de la nutria gigante (Pteronura brasiliensis) en la zona de influencia de Inírida (Bajo río Inírida) Guainía, Colombia PDF 211KB

Velasco, D.M. (2006). Investigación en Selva Mataven: Estudio preliminar de la población de perro de agua (Pteronura brasiliensis) en la zona de las comunidades de Giro y Morichal, resguardo Selva Matavén, Vichada, Colombia. PDF 460KB

Ecuador

Carrera, P. (2003). Solapameinto de nicho entre el hombre y la nutria gigante (Pteronura brasiliensis, Carnivora: Mustelidae) en la cuenca baja del Río Yasuní, Parque Nacional Yasuní, Amazonía Ecuatoriana.  Thesis for BSc, Departamento de Ciencias Biolólicas, Facultad de Ciencias Exactas y Naturales, Pontificia Universidad Católica del Ecuador
Part 1 PDF 3MB
Part 2 PDF 3MB
Part 3 PDF 3MB

Carrera-Ubidia, P.M. (2006). Giant Otters ( Pteronura brasiliensis) and Humans in the Lower Yasuní Basin,
Ecuador: Spacio-temporal Activity Patterns and Their Relevance for Conservation. Thesis for MSc in Wildlife Science, Oregon State University, Oregon, USA.
PDF 5.5MB

Fersen, L. von (1997). The Giant Otter in Ecuador. IUCN Otter Spec. Group Bull. 14 (1) April 1997; pp. 20-23

Melendres, A.P. (1978). Notas sobre las nutrias de río del Ecuador. In: Proceedings First Working Meetting of the Otter Specialist Group, Paramaribo, Suriname, 1977, N. Duplaix (ed.); pp. 56-66

Tirira, D. (Ed.) (2001). Libro rojo de los mamíferos del Ecuador. SIMBIOE/EcoCiencia/Ministerio del Ambiente/UICN. Serie Libros Rojos del Ecuador, Tomo 1. Publicación Especial sobre los Mamíferos del Ecuador4. Quito; 236 pp.

Utreras, V. And I. Araya. (2002). Distribution And Conservation Status Of The Neotropical Otter (Lutra longicaudis) And Giant Otter (Pteronura brasiliensis) In Ecuador. Pp. 365-369 In: Dulfer, R., Conroy, J. Nel & A. C. Gutleb (Eds.). Proccedings VIIth International Otter Colloquium. Trebon. IUCN Otter Specialist Group Bulletin. Volume 19A, Special Issue.

Utreras, V., Araya, I., Denkinger, J. and Rodriguez, M. (1997). Study on the Ecology and Ethological Aspects of the Giant Otter Pteronura brasiliensis (Carnivora: Mustelidae) in the Parque Nacional Yasuní, Ecuador. IUCN/SSC Otter Specialist Group Bulletin 14 (1): 20 - 23

Utreras, V. & J. P. Jorgenson. (2003). Aspectos Sobre La Cacería Y La Distribución Actual E Histórica De La Nutria Gigante (Pteronura Brasiliensis) En La Amazonía Ecuatoriana. Pp. 130-135 En: Polanco-Ochoa, R. (Ed.). 2003. Manejo De Fauna Silvestre En Amazonía Y Latinoamérica. Selección De Trabajos V Congreso Internacional. CITES, Fundación Natura. Bogotá, Colombia. 446 Pp.

Utreras, V. And L. Pinos. (2003). Camera Trap Use For Studying Giant Otters (Pteronura Brasiliensis) In The Yasuní Biosphere Reserve, Ecuadorian Amazon. IUCN Otter Specialist Group Bulletin, 20 (2): 69-71.

Utreras, V., E. Suárez, G. Zapata-Ríos, G. Lasso and L. Pinos. (2005). Dry and Rainy Season Estimations of Giant Otter, Pteronura brasiliensis, Home-Range in the Yasuní National Park, Ecuador. The Latin American Journal of Aquatic Mammals 4 (2): 1-4. PDF 182KB

Utreras, V. Y D. Tirira. (2001). Nutria Gigante (Pteronura brasiliensis). Pp. 61-63 En: D. Tirira (Ed.), Libro Rojo De Los Mamíferos Del Ecuador. SIMBIOE/Ecociencia/Ministerio Del Ambiente/UICN. Serie Libros Rojos Del Ecuador, Tomo 1. Publicación Especial Sobre Los Mamíferos Del Ecuador 4. Quito.

French Guiana

Jacques, H. and Pelsy, C. (2004). Les Loutres Géantes en Guyane Française, premiers résultats (distribution, impact de l'orpaillage) in Jacques H., Leblanc F. & Moutou F. (2005) Conservation de la Loutre. Société Française pour l'Etude et la Protection des Mammifères, Groupe Mammalogique et Herpétologique du Limousin. Paris, Limoges, 200 p. PDF 109KB

Jacques, H., Pelsy, C., de Thoisy, B. (2000) La loutre géante en Guyane Française : Rapport de mission sur le Litani

Jacques, H., Pelsy, C (2000) La loutre géante en Guyane Française : Rapport de mission sur l’Arataï

de Thoisy, B.,  Catzeflis, F., Richard-Hansen, C.,  Lavergne, A. &  Delcourt, B. (2006). Tapir De Tierras Bajas Y Nutria Gigante : Especies Focales De Los Ecosistemas Amenazados En Guayana Francesa. VII Congresso Internacional Sobre Manejo De Fauna Silvestre Na Amazônia E América Latina. Ilhéus, Bahia, Brasil.

Guyana

Asso. Kwata. (2006). Suivi des populations chez la loutre géante: aide à la conservation d'une espèce emblématique des cours d'eau amazoniens. Unpublished report, Kwata / Conseil Régional de la Guyane

Asso. Kwata. (2005). Ecologie moléculaire et génétique des populations chez la loutre géante: outils d'aide à la conservation d'une espèce emblématique des cours d'eau amazoniens. Unpublished report, Kwata / Direction Régionale de l'Environnement de la Guyane.

Barnett, A., Shapley, R., Lehman, S., Henry, E. & Benjamin, P. (2000). Records of the Giant Otter, Pteronura brasiliensis, from Guyana (Final version). IUCN Otter Spec. Group Bull. 17 (2) 2000; pp. 65-74

Laidler, E. (1982). A Study of the Giant Otter Pteronura brasiliensis - Movement Patterns, Territoriality, Development and Ecology in Guyana. Otters, Journal Otter Trust, 1982 Vol. 1 (6); pp. 13-21

Laidler, P.E. (1984). The behavioural ecology of the giant otter in Guyana. Doctoral Dissertation, Univ. of Cambridge 1984; 319 pp. (13581, D58408-85)

Shackley, M. (1998). Designating a Protected Area at Karanambu Ranch, Rupununi Savannah, Guyana: Resource Management and Indigenous Communities. Ambio Vol. 27 No. 3, May 1998; pp. 207-210

Shackley, M. (1996). Giant Otters at Karanambu. Unpublished; 13 pp.

Shackley, M. (1996). Feasibility Study for a Protected Area at Karanambu Ranch, Guyana; 88 pp.

Peru

Brack Egg, A. (1977). Situacion Actual de las Nutrias (Lutrinae, Mustelidae) en el Peru. In: Proceedings First Working Meeting of the Otter Specialist Group, Paramaribo, Suriname, 1977, N. Duplaix (ed.); pp. 76-84

Brecht-Munn, M. (1985). Lobo grande de río. Reporte Manu. Centro de Datos para la conservacion, Univ. Nacional Agraria, Lima, Peru; 6 pp.

Frank K. (2000). Análisis de un modelo para el manejo del Lobo de Río en el Parque Nacional del Manú. In: Lobo de Río, Resumen del Simposio 1998. P. 10-20. Inrena150141-2000-0127. Lima, Peru.

Grimwood, I. (1969). Giant Otter (Pteronura brasiliensis). In Endangered Mammals in Peru. Oryx Vol. IX; pp. 416

Groenendijk, J. (1998). De Reus van Zuid-Amerika. Ecologie en Ontwikkeling, Jaargang 6, No 1, 1998; pp. 15 – 17. Nieuwsbrief van het Nederlands Comité voor IUCN

Groenendijk, J. and Hajek, F. (2006). Giants of the Madre de Dios. Ayuda para Vida Silvestre Amenazada-Sociedad Zoologica de Francfort Peru. ISBN: 9972-2585-1-3, pp. 160.

Groenendijk, J. & Hajek, F. (2001). Project Report 2000 - Internal Report of the Frankfurt Zoological Society Giant Otter Conservation Project

Groenendijk, J. & Hajek, F. (2000). Project Report 1999 – Internal Report of the Frankfurt Zoological Society Giant Otter Conservation Project

Groenendijk, J. & Hajek, F. (1997). Las Piedras '96: an expedition to the Las Piedras river, south-eastern Peru. Produced by the Netherlands Committee for IUCN; pp. 44

Groenendijk, J., Hajek, F., Isola, S. & Schenck, C. (2001). Giant Otter Project in Peru: Field Trip and Activity Report, 2000. IUCN Otter Spec. Group Bull. 18(2): 76 - 85

Groenendijk, J., Hajek, F., Isola, S. & Schenck, C. (2000). Giant Otter Project in Peru: Field Trip and Activity Report, 1999. IUCN Otter Spec. Group Bull. 17(1)2000; pp. 34 - 45

Groenendijk, J., Hajek, F., Schenck, C. & Staib, E. (2001). Monitoreo del Lobo de Río (Pteronura brasiliensis) en la Reserva de Biosfera del Manu : metodologias y resultados. In: El Manu y Otras Experiencias de Investigacion y Manejo de Bosques Tropicales, Symposium Internacional, Puerto Maldonado, Peru, 4-7 June 2001

Groenendijk, J., Hajek, F., Schenck, C. & Staib, E. (in prep.). A Decade of Giant Otter (Pteronura brasiliensis) Population Dynamics in the Manu Biosphere Reserve, Peru. Paper presented at the VIII International Otter Colloquium, Valdivia, Chile, 19-27 January, 2001

Hajek, F. & Groenendijk, J. (in prep.). Propuesta para un Plan de Manejo de Habitats Acuáticos de la Reserva de Biosfera del Manu en base a monitoreo de especies indicadores; 37 pp.

Hajek, F. and Groenendijk, J. (2006). Lobos del Rio Madre de Dios. Ayuda para Vida Silvestre Amenazada-Sociedad Zoologica de Francfort Peru. ISBN: 9972-2585-2-1, pp. 160

Hajek, F. & Groenendijk, J. (2001). Informe Annual 2000 – Proyecto Lobo de Río. Report of the Frankfurt Zoological Society Giant Otter Conservation Project, presented to the Instituto Nacional de Recursos Naturales (INRENA)

Hajek, F. & Groenendijk, J. (2000). Informe Annual 1999 – Proyecto Lobo de Río. Report of the Frankfurt Zoological Society Giant Otter Conservation Project, presented to the Instituto Nacional de Recursos Naturales (INRENA)

Hajek, F., Groenendijk, J., Schenck, C. & Staib, E. (2001). El Turismo de Naturaleza y Manejo de Habitat de Lobo de Río (Pteronura brasiliensis) en el Sureste del Perú. In: El Manu y Otras Experiencias de Investigacion y Manejo de Bosques Tropicales, Symposium Internacional, Puerto Maldonado, Peru, 4-7 June 2001

Isola S. (1999). Evaluación del estado de conservación de tres especies de mamiferos dulce aquicolas sudamericanos: Lobo de rio (Pteronura brasiliensis), delfín rosado (Inia geoffrensis) y Manatí (Trichechus inunguis). Report WWF: 103.

Isola, S. (2000). Determinacion de la distribucion y abundancia de Lobo de Rio (Pteronura brasiliensis) en la Reserva Nacional Pacaya Samiria. Tesis para optar el Grado de Ingeniero Forestal, Univ. Nac. Agraria la Molina, 80 pp.

Isola, S. & Benavides, J. (2001). El Lobo de Rio: una especie bandera para la Reserva Comunal Tamshiyacu Tahuayo. Internal Report.

Khanmoradi, H. (1994). Untersuchungen zur Nahrungsökologie der Riesenotter (Pteronura brasiliensis) in Peru. Diplom-Arbeit, Uni. München; 121 pp.

Llosa T. (1997). Lobos del Rio en el Rio Candamo, Puno, Peru. Internal Report Conservation International: 20

Schenck, C. (1999). “Lobo de Río (Pteronura brasiliensis) – Presencia, uso del hábitat y protección en el Perú. Spanish translation of German PhD dissertation : Vorkommen, Habitatnutzung und Schutz des Riesenotters (Pteronura brasiliensis) in Peru (1996), Munich

Schenck, C. (1996). Vorkommen, Habitatnutzung und Schutz des Riesenotters (Pteronura brasiliensis) in Peru. Diss. Univ. Munchen, ISBN 3-8265-3126-4; 180 pp.

Schenck, C. (1995). Status, Habitat Use and Conservation of Giant Otters (Pteronura brasiliensis) in Peru. Habitat 11 - Proceedings VI International Otter Colloquium, Pietermaritzburg 1993. Reuther, C. & Rowe-Rowe, D. (eds.); pp. 69-71

Schenck, C. & Staib, E. (1995). The Giant Otter Project in Peru 1995. IUCN Otter Spec. Group Bull. 12 / October 1995; pp. 25-30

Schenck, C. & Staib, E. (1995). News from the Giant Otter Project in Peru. IUCN Otter Spec. Group Bull. 11 / April 1995; pp. 5-8

Schenck, C. & Staib, E. (1993). Giant Otter Study in Peru Completed. IUCN Otter Spec. Group Bull. 8/1993; pp. 44

Schenck, C. & Staib, E. (1992). Giant Otters in Peru. IUCN Otter Spec. Group Bull. 7/1992; pp. 24-26

Schenck, C., Estremadoyro, F. & Staib, E. Primeras evaluaciones para determinar la existencia de lobo de rio (Pteronura brasiliensis) en el ambito de la Reserva Nacional Pacaya-Samiria en el Peru. Unpublished; 11 pp.

Schenck, C., Groenendijk, J. & Hajek, F. (1999). Giant Otter Project in Peru – Field Trip and Activity Report, 1998. IUCN Otter Spec. Group Bull. 16(1)1999; pp. 33-43

Schenck, C., Staib, E. & Storch, I. (1997). 1996 News from the Giant Otter Project in Peru. IUCN Otter Spec. Group Bull. 14 (1) April 1997; pp. 13-19

Schweizer J. (1986). Ocorrencia e alguns dados biológicos de ariranha (Pteronura brasiliensis, Mustelidae) no Rio Negro. 20 Reuniao de trabalho de especialistas en mamiferos aquaticos da America do Sul. Resumos: 48.

Staib, E. (1995). Social behavior and ecology of the giant otter (Pteronura brasiliensis) in Peru. In: Habitat Arbeitsberichte der Aktion Fischotterschutz e.V.. Proceedings VI. International Otter Colloquium Pietermaritzburg 1993 (C. Reuther and D. Rowe-Rowe, eds.). GN-Gruppe Naturschutz GmbH: Hankensbüttel.

Staib, E. (1993). Social Behaviour and Ecology of the Giant Otter (Pteronura brasiliensis) in Peru. Habitat 11 - Proceedings VI International Otter Colloquium, Pietermaritzburg. Reuther, C. & Rowe-Rowe, D. (eds.); pp. 72-74

Staib, E. (submitted 2001). Öko-Ethologie von Riesenottern (Pteronura brasiliensis) in Peru, PhD dissertation, Munich University, Germany.

TReeS (Tambopata Reserve Society) (1994). Report of the TReeS Tambopata-Candamo Expedition. A biological survey in the Tambopata-Candamo Reserved Zone South-East Peru (12.08.1992-20.09.1992). London, England. 171 pp.

Suriname

Duplaix, N. (1978). Synopsis of the status and ecology of the giant otter in Suriname. In: Proceedings First Working Meeting of the Otter Specialist Group, Paramaribo, Suriname, 1977, N. Duplaix (ed.); pp. 48-54

Duplaix, N. (1979) Giant otters in Suriname. Oryx. 14(4):439

Duplaix, N. (1980). Observations on the Ecology and Behavior of the Giant River Otter (Pteronura brasiliensis) in Suriname. Rev. Ecol. (Terre Vie), vol. 34 (1980); pp. 496-620 PDF 6MB

Duplaix, N. (1982) Contribution à l'Ecologie et à l'Ethologie de Pteronura brasiliensis (Carnivora, Lutrinae): Implications Evolutives. Doctorat Thesis, Univ. of Paris-Sud, 352 pp.

Duplaix, N., M. Lingaard, & C. Sakimin (2001) A Survey of Kaburi Creek, West Suriname and its Conservation Implications 2001, Oceanic Society, Oceanic Society, 34 pp

Venezuela

Mondolfi, E. & Trebbau, P. (1997). Distribution and Status of the Giant Otter (Pteronura brasiliensis) in Venezuela. Second report, Nov. 1997, unpublished; 6 pp.

Mondolfi, E. & Trebbau, P. (1978). Distribution and Status of the Giant Otter (Pteronura brasiliensis) in Venezuela. In: Proceedings First Working Meeting of the Otter Specialist Group, Paramaribo, Suriname, 1977, N. Duplaix (ed.); pp. 44-47

Ecology and Behavior

Carter, S.K. & Rosas, F.C.W. (1997). Biology and conservation of the Giant Otter Pteronura brasiliensis. Mammal Rev. 1997, Vol. 27, No.1, pp. 1-26 PDF 6MB

Duplaix, N. (1980). Observations on the Ecology and Behavior of the Giant River Otter (Pteronura brasiliensis) in Suriname. Rev. Ecol. (Terre Vie), vol. 34 (1980); pp. 496-620

Duplaix, N. (1982) Contribution à l'Ecologie et à l'Ethologie de Pteronura brasiliensis (Carnivora, Lutrinae): Implications Evolutives. Doctorat Thesis, Univ. of Paris-Sud, 352 pp.

Evangelista, E., (2004). Change of partners in a Giant otter Alpha couple. IUCN Otter Spec. Group Bull. 21(1): 47-51

Gómez, J.R. (1999). Ecología alimentaria de la Nutria Gigante (Pteronura brasiliensis) en el bajo Río Bita (Vichada, Colombia). Tesis, Universidad Javeriana, Santafé de Bogotá, Colombia.

Laidler, E. (1982). A Study of the Giant Otter Pteronura brasiliensis - Movement Patterns, Territoriality, Development and Ecology in Guyana. Otters, Journal Otter Trust, 1982 Vol.1 (6); pp. 13-21

Laidler, P.E. (1984). The behavioural ecology of the giant otter in Guyana. Doctoral Dissertation, Univ. of Cambridge 1984; 319 pp. (13581, D58408-85)

Mourao, G. & Carvalho,L. (2001) Cannibalism among Giant Otters (Pteronura brasiliensis) Mammalia 65, n°2, 2001 : 225-227

Ribas, C. and G. Mourao (2004). Intraspecific agonism between Giant otter groups PDF 614KB

Rosas, F.C.W., Zuanon, J.A.S. & Carter, S.K. (1999). Feeding Ecology of the Giant Otter, Pteronura brasiliensis. BIOTROPICA 31(3); pp. 502-506

Schenck, C., Staib, E. & Yasseri, A.M. (1995). Unterwasserlaute bei Riesenottern (Pteronura brasiliensis). Z. Saugetierkunde 60; pp. 310-313

Schenck, C. (1997).Vorkommen, Habitatnutzung und Schutz des Riesenotters (Pteronura brasiliensis) in Peru. Shaker Verlag, München.

Staib, E. 1995. Social behavior and ecology of the giant otter (Pteronura brasiliensis) in Peru. In: Habitat Arbeitsberichte der Aktion Fischotterschutz e.V.. Proceedings VI. International Otter Colloquium Pietermaritzburg 1993 (C. Reuther and D. Rowe-Rowe, eds.). GN-Gruppe Naturschutz GmbH: Hankensbuttel.

Schroeter, W. (1982). Verhaltensbeobachtungen am Riesenotter (Pteronura brasiliensis, Gmelin, 1788) in Gefangenschaft. Saugetierkundliche Mitt. 30(3) 1982; pp. 216-220

Staib, E. (1993). Social Behaviour and Ecology of the Giant Otter (Pteronura brasiliensis) in Peru. Habitat 11 - Proceedings VI International Otter Colloquium, Pietermaritzburg. Reuther, C. & Rowe-Rowe, D. (eds.); pp. 72-74

Staib, E. (submitted 2001). Öko-Ethologie von Riesenottern (Pteronura brasiliensis) in Peru, PhD dissertation, Munich University, Germany.

Trebbau, P. (1978). Some observations on the Mating Behaviour of the Brazilian Giant Otter (Pteronura brasiliensis). Zool. Garten N. F., Jena 48 (1978) 2/3, S.; pp. 187-188

Utreras, V., Araya, I., Denkinger, J. and Rodriguez, M. (1997). Study on the Ecology and Ethological Aspects of the Giant Otter Pteronura brasiliensis (Carnivora: Mustelidae) in the Parque Nacional Yasuní, Ecuador. IUCN/SSC Otter Specialist Group Bulletin 14 (1): 20 - 23

Status and Conservation

Carter, S.K. & Rosas, F.C.W. (1997). Biology and conservation of the Giant Otter Pteronura brasiliensis. Mammal Rev. 1997, Vol. 27, No.1, pp. 1-26 PDF 6MB

Barnett, A., Shapley, R., Lehman, S., Henry, E. & Benjamin, P. (2000). Records of the Giant Otter, Pteronura brasiliensis, from Guyana (Final version). IUCN Otter Spec. Group Bull. 17(2) 2000; pp. 65-74

Chehebar, C. (1989). The Status of Otters in Latin America. Habitat 6 - Proceedings V Internat. Otter Colloquium, Hankensbuttel 1989. Reuther, C. & Rochert, R. (eds.); pp. 9-14

Duplaix, N. (1980). Observations on the Ecology and Behaviour of the Giant River Otter (Pteronura brasiliensis) in Suriname. Rev. Ecol. (Terre Vie), vol. 34 (1980); pp. 496-620

Duplaix, N. (1980). Giant otters : "Big water dogs in peril". National Geographic Magazine 158(1) 1980; pp. 130-142 PDF 2.5MB

Duplaix, N. (2001). Evaluation of the Animal and Plant Trade in the Guianas: Preliminary Findings. WWF-Guianas PDF 1MB

Duplaix, N. (2002). Giant Otter, Final Report. WWF-Guianas Rapid River Bio-assessments and Giant Otter Conservation Project - FG-40 FY2002 PDF 8MB

Duplaix, N. & J.A. Davis (1983) Otter, Identification Manual for Customs Officials. 30; Secretariat of the Convention on International Trade in Endangered Species of Wild Fauna and Flora. (CITES), Gland, Switzerland.

Ergueta, P. & Morales, C. de [eds.] (1996). Libro rojo de los vertebrados de Bolivia. CDC - Bolivia; pp. 171-172

Fonseca, G. & Rylands, A. & Costa, C. & Machado, R. & Leite, Y. (1994) Livro vermelho dos Mamiferos brasileiros ameaçados de extinçao Belo Horizonte

Frank K. (2000). Análisis de un modelo para el manejo del Lobo de Río en el Parque Nacional del Manú. In: Lobo de Río, Resumen del Simposio 1998. P. 10-20. Inrena150141-2000-0127. Lima, Peru.

Groenendijk, J. (1998). A Review of the Distribution and Conservation Status of the Giant Otter (Pteronura brasiliensis), with Special Emphasis on the Guyana Shield Region. Commissioned by the International Fund for Animal Welfare, produced by the Netherlands Committee for IUCN, March 1998; 55pp.

Groenendijk J., F. Hajek, N. Duplaix, et al. (2005). Surveying and Monitoring Distribution and Population trends of the Giant Otter (Pteronura brasiliensis) Habitat 16: 1-53 PDF 7MB

Groenendijk, J., Hajek, F. & Schenck, C. (2004). Pteronura brasiliensis. In: IUCN 2004. 2004 IUCN Red List of
Threatened Species. <www.redlist.org>. Downloaded on 07 November 2005.
PDF 97KB

Harris, C. J. A. (1968). Otters,A Study of the Recent Lutrinae. London: Weidenfeld and Nicholson.

Hilton-Taylor C. (2000). 2000 IUCN Red List of Threatened Species. IUCN, Gland, Switzerland and Cambridge, UK.

INRENA (1999). Resumen del Simposio Internacional Sobre Conservacion del Lobo de Rio (Pteronura brasiliensis), Lima, 10-11 setiembre 1998. Produced by the Instituto Nacional de Recursos Naturales, Peru

IUCN (1996). Red List of Threatened Animals. Compiled and edited by Jonathon Baillie and Brian Groombridge, World Conservation Monitoring Centre. Published by IUCN and Conservation International; 448 pp.

Olson, D.M., Isola, S.E., Cogliano, M., Thieme, M.L. & Mobley, M. (2001). Designing a representative network of protected areas for giant otters (Pteronura brasiliensis) and other Neotropical freshwater megafauna. WWF-US Report.

Rosas, F. C. W. (2004)  Ariranha, Pteronura brasiliensis (Carnivora:Mustelidae). In: Renato Cintra. (Org.). História Natural, ecologia e conservação de algumas espécies de plantas e animais da Amazônia. Manaus: Editora da Universidade Federal do Amazonas (EDUA), 2004, p. 265-269. PDF 1.2MB

Rosas, F. C. W., Colares, E. P., Colares, I. G.& Silva, V. M. F. (1991). Mamíferos Aquáticos da Amazônia Brasileira. In: Adalberto Luis Val; Roberto Figliuolo; Eliana Feldberg. (Org.). Bases Científicas para Estratégias de Preservação e Desenvolvimento da Amazônia: Fatos e Perspectivas. 1 ed. Manaus: Imprensa Universitária, 1991, v. 1, p. 405-411.

Rosas, F. C. W., Waldemarin, H. & Mattos, G. E. (2006)   Ariranha, Pteronura brasiliensis (Zimmermann, 1780) (in press). In: A. B.Machado; G. Drummond; A. Paglia. (Org.). Livro Vermelho da Fauna Brasileira Ameaçada de Extinção. Belo Horizonte: Fundação Biodiversitas, 2006.

Schenck, C. and Staib, E. 1998. Status, habitat use and conservation of giant otters in Peru In: DUNSTONE N, GORMAN M.: Behaviour and Ecology of Riparian Mammals, Cambridge University Press, 359-370.

Staib, E. and Schenck, C. 1994. Giant Otter…A giant under even bigger pressure. Wildbiologische Gesellschaft, Munich and Frankfurt Zoological Society, Frankfurt.

Tirira, D. (Ed.) (2001). Libro rojo de los mamíferos del Ecuador. SIMBIOE/EcoCiencia/Ministerio del Ambiente/UICN. Serie Libros Rojos del Ecuador, Tomo 1. Publicación Especial sobre los Mamíferos del Ecuador4. Quito; 236 pp.

Giant otters in Captivity

Autuori M.P. & Deutsch, L.A. (1997). Contribution to the Knowledge of the Giant Brazilian Otter, Pteronura brasiliensis (Gmelin 1788), Carnivora, Mustelidae - Rearing in Captivity. Zool. Garten N.F., 47 (1997) 1, S.; pp. 1-8

Bartmann, W. 1996. International census of the giant river otter (P. brasiliensis) in captivity (1996). Tierpark Dortmund, Dortmund, Germany. Unpublished report.

Carter, S. K., Rosas, F. C. W., Cooper, A. & Duarte, A. C. C. (1999). Consumption rate, food preferences and transit time of captive giant otter, Pteronura brasiliensis: Implications for the study of wild populations.. Aquatic Mammals, v. 25, n. 2, p. 79-90, 1999.

Corredor, G. & Tigreros Muñoz, N. (2006). Reproduction, behaviour and biology of the Giant river otter Pteronura brasiliensis at Cali Zoo. Int. Zoo Yb. 40: 360–371 PDF 644KB

Duplaix-Hall, N. (1972). Notes on maintaining river otters in captivity. Inter. Zoo Yrbk. 12:178-181.

Duplaix - Hall, N. (1975). River otters in captivity: a review. In Martin, R.D. (ed.), Breeding Endangered Species in Captivity, Academic Press, London; pp. 315-327

Duplaix, N. (1982) Contribution à l'Ecologie et à l'Ethologie de Pteronura brasiliensis (Carnivora, Lutrinae): Implications Evolutives. Doctorat Thesis, Univ. of Paris-Sud, 352 pp.

Flügger, M. von (1997). Haltung, Zucht und Erkrankungen von Riesenottern (Pteronura brasiliensis) in Carl Hagenbecks Tierpark. Verh. ber. Erkrg. Zootiere (1997) 38; pp. 125-130

Foster-Turley, P. 1990. Otters in Captivity. In: Otters: An Action Plan for their Conservation (Foster-Turley, P., Macdonald, S. and Mason, C., eds.). International Union for Conservation of Nature.

“Genealogical Meeting on Giant Otters in Captivity in Brazil”, lectures/workshop discussions, December 3-4, 1998. Jardim Zoologico de Brasília , Brasilia, Brazil. Daniel Louzada da Silva and Dr. Claudio Valladares-Padua, conference organizers.

Hagenbeck, C. and Wünnemann, K. 1992. Breeding the giant otter Pteronura brasiliensis at Carl Hagenbeck’s Tierpark. Int. Zoo Yrbk. 31:240-245.

Gómez, J.R., Jorgenson, J. & Valbuena, R. (1999). Report on the Rehabilitation and Release of Two Giant River Otter (Pteronura brasiliensis) Pups in the Bita River (Vichada, Colombia). IUCN Otter Spec. Group Bull. 16(2)1999; pp. 86-89

Hagenbeck, C. & Wünnemann, K. (1992). Breeding the Giant Otter (Pteronura brasiliensis) at Carl Hagenbecks Tierpark. New Developments in the Zoo World, Internat. Zoo Yb. (1992) 31; pp. 240-245

Kranz, K. (Oct. 1996). Suggested Design Criteria for Giant Otter Enclosures.; 4 pp.

Marsicano, G., Marmontel, M., Rosas, F. C. W. & Souza, R. H. S. Contenção de lontras com Cloridrato de Ketamina.. A Hora Veterinária, Porto Alegre, v. 6, n. 31, p. 5-8, 1986. PDF 1MB

McTurk, D. and L. Spelman. (2005) Hand-rearing and rehabilitation of orphaned Giant otters Pteronura brasiliensis, on the Rupununi River, Guyana, South America. Zoo Biol. 24:153-167 (2005), 15 pages PDF 130KB

Nehring, A. (1901) EinigeNotzen uber die Lutra (Pteronura) paranensis des niesigen Zoologoscjer Garten. Sber. Ges. Naturf. Freude Berlin, 1901: 133-135.

Rosas, F. C. W., Rocha, C. S. & Mattos, G. E. (2006). A method to measure captive giant otters (Pteronura brasiliensis) without sedation. The Latin American Journal of Aquatic Mammals, v. 5, p. 65-66, 2006. PDF 337KB

Rocha, C. S., Rosas, F. C. W., Mattos, G. E. & Lazzarini, S. M. (2005). Body weight-length relationships of giant otters (Pteronura brasiliensis). In: XVI Biennial Conference on the Biology of Marine Mammals, 2005, San Diego, CA - USA. Livro de resumos da XVI Biennial Conference on the Biology of Marine Mammals, 2005.

Salvo Souza, R. and Best, R. 1982. Unpublished Report. Contribuicao sobre o comportamento da ariranha (Pteronura brasiliensis Gmelin 1788) em cativeiro. Mamiferos Aquaticos INPA, Manus. AM. Contribuicao no. 34 da Divisao de Mamiferos Aquaticos, INPA Manaus.

Schenck, C., Staib, E. & Yasseri, A.M. (1995). Unterwasserlaute bei Riesenottern (Pteronura brasiliensis). Z. Saugetierkunde 60; pp. 310-313

Skeldon, P.C. (1961). Exhibiting Aquatic Mammals - Brazilian Giant Otters (Pteronura brasiliensis) at Toledo Zoo. Internat. Zoo Yb. Vol.3, 1961; pp. 30-31

Sykes, S. 1998. “Historical census, management, and rearing success of giant otters (P. brasiliensis) in zoos: Need for an international studbook.” Lecture for “Genealogical Meeting on Giant Otters in Captivity in Brazil”, Jardim Zoologico de Brasília, Brazil: Dec. 2-4,1998.

Sykes, S. 1997-1999. “South American Giant Otter (Pteronura brasiliensis) Husbandry, Management, and Historical Census Survey.” Zoological Society of Philadelphia, PA, USA. Unpublished report.

Sykes-Gatz, S. 2000. “Giant otters (P. brasiliensis) in zoos: An overview and need for an international studbook.” Lecture for “Symposium-Haltung und Zucht des Riesenotters (P. brasiliensis)”, 17.06.00. Dortmund Zoo, Germany.

Sykes-Gatz, S. (2001). Husbandry and Management of the Giant Otter (Pteronura brasiliensis). Münster: Schüling, 2001, ISBN 3-930962-03-9; 30 pp.

Sykes-Gatz, S. (2001). Pteronura brasiliensis in Captivity/Studbook: An International Historical Overview and Management Recommendations. For: Otters: An Action Plan for their Conservation (2001). International Union for Conservation of Nature (IUCN), Otter Specialist Group, Otter in Captivity, Sub-chapter 2.6.11.

Sykes-Gatz, S. (2005). International Giant Otter Studbook Husbandry and Management Information and Guidelines. Husbandry and Management of the Giant Otter (Pteronura brasiliensis), 2nd Edition. Zoo Dortmund, Dortmund, Germany, Pp. 276. PDF 4MB

Sykes-Gatz, S. & Gatz, V. (2005). Photographs for the International Giant Otter Studbook Husbandry and Management Information and Guidelines (2005). Zoo Dortmund, Dortmund, Germany. PDF 5MB

Sykes-Gatz, S. & Gatz, V.( 2004). International Studbook for the Giant Otter (Pteronura brasiliensis) (Gray 1867), 1st Edition 2004, Zoo Dortmund, Dortmund, Germany. PDF 237KB

Symposium - Haltung und Zucht des Riesenotters (Pteronura brasiliensis), lectures/discussions, June 17, 2000. Dortmund Zoo, Dortmund, Germany. Dr. Wolf Bartmann, conference organizer.

Trebbau, P. 1972. Notes on the Brasilian giant otter (P. brasiliensis) in captivity. Zool. Garten 41:152-156.

Trebbau, P. (1978). Some observations on the Mating Behaviour of the Brazilian Giant Otter (Pteronura brasiliensis). Zool. Garten N. F., Jena 48 (1978) 2/3, S.; pp. 187-188

Wünnemann, K. 1995. Giant Otter Husbandry. Pp.181-184. In: Husbandry Handbook for Mustelids (J. Partridge, ed.). Association of British Wild Animal Keepers.

Wünnemann, K. (1995). Breeding Giant Otters (Pteronura brasiliensis) in Germany. Habitat 11 - Proceedings VI International Otter Colloquium, Pietermaritzburg. Reuther, C. & Rowe-Rowe, D. (eds.); pp. 75-77

Zeller, F. (1960). Notes on the Giant Otter (Pteronura brasiliensis Zimm) at Cologne Zoo. Internat. Zoo Yb. Vol.2, 1960; pp. 81

Useful Captive Husbandry Information

Baker, A.J.; Baker, A.M. and Thompson, K.V. 1996. Parental Care in Captive Mammals. Pp.497- 512. In: Wild Mammals in Captivity - Principles and Techniques (D.G. Kleiman, M.E. Allen, K.V. Thompson and S. Lumpkin, eds.). University of Chicago Press: Chicago.

B.M.E.L.F. (Bundesministerium für Ernährung, Landwirtschaft und Forsten (Hrsg.) 1996. Mindestanforderungen an die Haltung von Säugetieren. Bonn, Deutschland.

Crissey, S.D. 1998. Handling Fish Fed to Fish-Eating Animals: A Manual of Standard Operating Procedures. U.S. Department of Agriculture, Agriculture Research Service, National Agricultural Library.

Hancocks, D. 1980. Bringing nature into the zoo: Inexpensive solutions for zoo environments. International Journal for the Study of Animal Problems 1:170-177.

International Species Information System; ISIS Mammal Abstracts & ISIS Specimen Reports 1995-1997. (Data base via computer network.)

Lewis, J. C.M., 1995. Veterinary Considerations. Pp.203-221. In: Husbandry Handbook for Mustelids (J. Partridge, ed.). Association of British Wild Animal Keepers.

Merck Veterinary Manual. Sixth Edition, 1986. A Handboook of Diagnosis, Therapy, and Disease Prevention and Control for the Veterinarian. Merck & Co., Inc. Rahway, N.J. (now exists as a CD)

Worley, K. E. and Hare, V. J. 1995. An overview of the 1995 Environmental Enrichment Conference. The Shape of Enrichment, vol. 4, No. 4, November.

Taxonomy, Anatomy and Phylogenetics

Anderson, S. (1997). Mammals of Bolivia, Taxonomy and Distribution. Bull. Am. Mus. Nat. Hist., No. 231; pp. 343 – 344

Asso. Kwata. (2005). Ecologie moléculaire et génétique des populations chez la loutre géante: outils d'aide à la conservation d'une espèce emblématique des cours d'eau amazoniens. Unpublished report, Kwata / Direction Régionale de l'Environnement de la Guyane.

Carter, S.K., Rosas, F.C.W., Cooper, A.B. & Cordeiro-Duarte, A.C. (1999). Consumption Rate, Food Preferences and Transit Time of Captive Giant Otters Pteronura brasiliensis: Implications for the Study of Wild Populations. Aquatic Mammals 25 (2): 79 - 90 PDF 2.3MB

Duplaix, N. (1982) Contribution a l'Ecologie at a l'Ethologie de Pteronura brasiliensis (Carnivora, Lutrinae): Implications Evolutives. Doctorat Thesis, Univ. of Paris-Sud, 352 pp.

Duplaix, N. & J.A. Davis (1983) Otter, Identification Manual for Customs Officials. 30; Secretariat of the Convention on International Trade in Endangered Species of Wild Fauna and Flora. (CITES), Gland, Switzerland.

Dias Fonseca, F.R., Malm, O. & Francine Waldemarin, H.. (2005) Mercury Levels in tissues of Giant otters (Pteronura brasiliensis) from the Rio Negro, Pantanal, Brazil. Env. Res.98 (2005): 368-371 PDF 199KB

Harris, C. J. A. (1968). Otters,A Study of the Recent Lutrinae. London: Weidenfeld and Nicholson.

Heran, I. (1984). Comments on the age-conditioned changes of skull proportions in some species of Mustelidae. Lynx (Prague) 22 1984; pp. 11-14

Koepfli, K-P. & Wayne, R.K. (1998). Phylogenetic relationships of otters (Carnivora: Mustelidae) based on mitochondrial cytochrome b sequences. J. Zool., Lond. (1998) 246, pp. 401-416

Machado, G. V. ; Rosas, F. C. W. ; Lazzarini, S. M. (2002). O arco aórtico e seus ramos, diretos e indiretos, na ariranha (Pteronura brasiliensis Zimmermann, 1780 - Carnivora, Mustelidae).. Arquivos de Ciências Veterinárias e Zoologia da Unipar, Umuarama - PR, v. 5, n. 2, p. 285-285, 2002. PDF 1MB

Machado, G. V. ; Rosas, F. C. W. ; Lazzarini, S. M. (2002) Lobação pulmonar e divisões bronquiais na ariranha (Pteronura brasiliensis Zimmermann, 1780 - Carnivora, Mustelidae). Arquivos de Ciências Veterinárias e Zoologia da Unipar, Umuarama - PR, v. 5, n. 2, p. 289-289, 2002. PDF 1MB

Machado, G. V. ; Rosas, F. C. W. ; Lazzarini, S. M. (2002). Topografia do cone medular na ariranha da Amazônia (Pteronura brasiliensis Zimmermann, 1780 - Carnivora, Mustelidae). Arquivos de Ciências Veterinárias e Zoologia da Unipar, Umuarama - PR, v. 5, n. 2, p. 297-297, 2002. PDF 1MB

Machado, G. V. ; Rosas, F. C. W. ; Lazzarini, S. M. (2002). Sobre a formação da veia cava cranial na ariranha da Amazônia (Pteronura brasiliensis Zimmermann, 1780). Arquivos de Ciências Veterinárias e Zoologia da Unipar, Umuarama - PR, v. 5, n. 2, p. 293-293, 2002. PDF 1MB

Murmann, W. von & Hagenbeck, C. (1984). Steinbildungen in den Calices renales beim brasilianischen Riesenfischotter (Pteronura brasiliensis). Der Praktische Tierarzt, Sonderdruck 65. Jahrgang 1984, Heft 9, Seite 748

Pohle, H. (1920) Die Unterfamilie der Lutrinae. Arch. Naturgesch, 85 Ab. H. Heft 9: 1-247.

Radinsky, L. B. (1968). Evolution of somatic sensory specialisation in otter brains. J. Comp. Neurol., 134: 495-505

Radinsky, L. B. (1981) Evolution of skull shape in carnivores. 1. Representative modern carnivores. Biol. J. Linnean Soc. 15 (4): 369-388

Vieira, C. O. da C. (1952) Notas sobre os mamiferos obtidos pela expedicao do Instituto Butanta as Rio das Mortes e Serra do Roncador. Papeis Dept. Zool. S. Paulo 10:117.

Physiology, Biochemistry and Genetics

Colares, E.P. & Best, R.C. (1991). Blood parameters of Amazon otters (Lutra longicaudis, Pteronura brasiliensis) Carnivora, Mustelidae. Comp. Biochem. Physiol. Vol. 99A, No. 4; pp. 513-515

Dunstone N. (1998). Adaptations to the semi-aquatic habit and habitat. In: Behaviour and Ecology of Riparian Mammals. Dunstone N. and Gorman M. (1998). Symposia of the Zoological Society of London 71. Cambridge University Press. pp. 391.

Gutleb, A.C., Schenck, C. & Staib, E. (1994). Giant Otter (Pteronura brasiliensis) at Risk? Total Mercury and Methylmercury Levels in Fish and Otter Scats, Peru. Ambio Vol. 26 No. 8, Dec. 1997; pp. 511 - 514

Gutleb, A.C., Schenck, C. & Staib, E. (1993). Total Mercury and Methylmercury Levels in Fish from the Department Madre de Dios, Peru. IUCN Otter Spec. Group Bull. 8/1993; pp. 16-18

Kleinschmidt, T., Braunitzer, G. & Scheill, H.-G. (1989). Carnivora: the primary structure of the giant otter (Pteronura brasiliensis, Mustelidae) hemoglobin. Biol. Chem. Hoppe-Seyler 370(4) 1989; pp. 303-308

Leonards, P.E. G (1997) PCBs in Mustelids - analysis, food chain transfer and critical levels. Academic thesis. Vrije Universiteit, Amsterdam; 210 pp.

Scheill, H.-G. & Gunther, A. (1984). Elektrophoretischer Vergleich einiger Erythrocyten-Enzyme der Otter Lutra Lutra (L. 1758), Lutra canadensis (Schreber 1776) und Pteronura brasiliensis (Gmelin, 1788). Saügetierkundliche Mitt. 32(1) 1984; pp. 63-66

Veterinary Problems and Diseases

Lewis, J. C.M., 1995. Veterinary Considerations. Pp.203-221. In: Husbandry Handbook for Mustelids (J. Partridge, ed.). Association of British Wild Animal Keepers.

Pimentel, T.L., Reis, M.L. & Passerino, A.S.M. Mustelidae – Biology, Management and Medicine. In : Biology, Management & Medicine of South American Wild Animals, Fowler, M.E. & Cubas, Z.S. (eds.).

Schenck, C. and Staib, E., Storch I. (1997). Domestic animal disease risks for Peruvian Giant Otters (Pteronura brasiliensis) IUCN Veterinary Specialist Group Newsletter, No. 14, p. 7-8.

Giant Otters and people, tourism, public education

Carrera, P. (2003). Solapameinto de nicho entre el hombre y la nutria gigante (Pteronura brasiliensis, Carnivora: Mustelidae) en la cuenca baja del Río Yasuní, Parque Nacional Yasuní, Amazonía Ecuatoriana.  Thesis for BSc, Departamento de Ciencias Biolólicas, Facultad de Ciencias Exactas y Naturales, Pontificia Universidad Católica del Ecuador
Part 1 PDF 3MB
Part 2 PDF 3MB
Part 3 PDF 3MB

Carrera-Ubidia, P.M. (2006). Giant Otters ( Pteronura brasiliensis) and Humans in the Lower Yasuní Basin,
Ecuador: Spacio-temporal Activity Patterns and Their Relevance for Conservation. Thesis for MSc in Wildlife Science, Oregon State University, Oregon, USA.
PDF 5.5MB

Donadio, A. (1978). Some Comments on Otter Trade and Legislation in Colombia. In: Proceedings First Working Meeting of the IUCN Otter Specialist Group, Paramaribo, Suriname, March 1977, N. Duplaix (ed.); pp. 34-42

Duplaix, N. (1980). Giant otters: Big water dogs in peril. National Geographic Magazine 158(1) 1980; pp. 130-142 PDF 2.5MB

Gómez, J.R. & Jorgenson, J. (1999). An Overview to the Giant Otter – Fisherman Problem in the Orinoco Basin of Colombia. IUCN Otter Spec. Group Bull. 16(2)1999; pp. 90-96

Frankfurt Zoological Society Giant Otter Project (2000). Help Protect the Jungle Giant – Giant Otters, a Unique and Endangered Mammal. Leaflet for tourists, also produced in Spanish

Hajek, F., Groenendijk, J., Schenck, C. & Staib, E. (2001). Fundamento de un Plan de Manejo para el Lago Sandoval, con énfasis en el Lobo de Río (Pteronura brasiliensis). Version 1.4, presented to the Instituto Nacional de Recursos Naturales (INRENA); 7 pp.

Hajek, F., Groenendijk, J., Schenck, C. & Staib, E. (2001). El Turismo de Naturaleza y Manejo de Habitat de Lobo de Río (Pteronura brasiliensis) en el Sureste del Perú. In: El Manu y Otras Experiencias de Investigacion y Manejo de Bosques Tropicales, Symposium Internacional, Puerto Maldonado, Peru, 4-7 June 2001

Hajek, F., Groenendijk, J., Schenck, C. & Staib, E. (in prep.). Nature Tourism and Giant Otter (Pteronura brasiliensis) Habitat Management in Southeastern Peru. Paper presented at the VIII International Otter Colloquium, Valdivia, Chile, 19-27 January, 2001.

Hammond Ds, Gond V, De Thoisy B, Forget Pm, Dedijn B. (2007). Causes and consequences of a tropical forest gold rush in the Guiana Shield, South America. Ambio, 36, in press.

Schenck, C. & Staib, E. (1992/93). Look at the Giant Otters (Pteronura brasiliensis)....and give them a chance! (Leaflet aimed at tourists to encourage responsible behaviour in the presence of Giant Otters)

Schenck, C., Staib, E., Groenendijk, J. & Hajek, F. (2001). The ‘Pepe, the Giant Otter’ Colouring Book Activity and Drawing Competition as an Awareness-raising Tool for School Children. The Wetlands Ambassador: Education and Public Awareness Methodologies for Otter Conservation. Ando, M. & Sasaki, H. (eds.), Feb. 2001; pp. 46-47

Schenck, C., Staib, E., Groenendijk, J. & Hajek, F. (2001). Raising Awareness about, and Managing the Impact of Tourism in Giant Otter Habitats in South-eastern Peru. The Wetlands Ambassador: Education and Public Awareness Methodologies for Otter Conservation. Ando, M. & Sasaki, H. (eds.), Feb. 2001; pp. 44-46

Staib, E., Schenck, C. (1994). Giant Otters and Ecotourism in Peru. IUCN Otter Spec. Group Bull. 9 / March 1994; pp. 7-8

Staib, E., Schenck, C. (1994). Giant Otter... A giant under even bigger pressure. (Also Spanish version). Zoologische Gesellschaft Frankfurt / Wildbiologische Gesellschaft Munchen e.V.; 36 pp.

Staib E., Schenck C. (1996). Las Nutrias de la Selva - Quien sabe distinguirlas? - Faltblatt zu Fischottern in Peru, Ausgabe in Spanisch, Auflage 2000, Zoologische Gesellschaft Frankfurt, ProNaturaleza

Smith, N.J.H. (1981). Caimans, capaybaras, otters, manatees and man in Amazonia. Biol. Conserv. 19; pp. 177-187

IUCN Otter Specialist Group Publications and Giant Otter Bulletins

Cabral, M. M. M., Mattos, G. E. & Rosas, F. C. W. (2005) Registering giant otter birth during the first semester in Balbina lake. Friends of the Giant Otter, Soc. Zool Frankfurt/Lima, Peru, p. 3 - 4, 31 dez. 2005.

Foster-Turley, P., Macdonald, S. & Mason, C. (eds.) (1990). Otters - An Action Plan for their Conservation. IUCN / SSC Otter Spec. Group; 126 pp.

Groenendijk, J. (Ed.) (2002). Friends of the Giant Otter. Bulletin No.4 / January 2002

Groenendijk, J. (Ed.) (2001). Friends of the Giant Otter. Bulletin No.3 / June 2001

Groenendijk, J. (Ed.) (2000). Friends of the Giant Otter. Bulletin No.2 / December 2000

Groenendijk, J. (Ed.) (2000). Friends of the Giant Otter. Bulletin No.1 / July 2000

Popular articles and books

General

Brown, D.O. (1996). Giant Otters. Nature Conservancy, Vol. 46, No.3, May/June; pp. 24-29

Duplaix, N. (1984). Otters. The Encyclopaedia of Mammals - Edited by Dr. David Macdonald. Unwin Hyman Ltd.; pp. 125-129

Munn, C. (2005). Big, friendly giants. BBC Wildlife (June): 34-39 PDF 414KB

Guyana

Duplaix, N. (2003). A Chorus of Otters. Whale (Journal of the Oceanic Society) 8: 8-12 PDF 1.5MB

Hardman, C. (2004). Caretaker of Guyana's Water Dogs. Américas 56 (2): 14-19 PDF 1.5MB

Laidler, L. (1986). Watching over the waterdogs. BBC Wildlife 4(4) 1986; pp. 168-171

Laidler, L. (1982). Bring back the giant. New Scientist 95 No. 1324, 1982; pp. 825-828

Laidler, K. & Laidler, L. (June 1995). Giant among Otters. Wildlife Conservation May / June 1995; pp. 42-47 & 75

Laidler, K. & Laidler, L. (1983). The River Wolf. Allen & Unwin, London 1983; 178 pp.

Laidler, K. & Laidler, L. (1981). The river wolf plays rough. International Wildlife 6(5) 1981; pp. 46-49

 Peru

Brecht-Munn, M. & Munn, C. (1988). The Amazon's Gregarious Giant Otters. Animal Kingdom Sept. / Oct. 1988; pp. 34-41

Schenck, C. & Staib, E. (1994). Die Wölfe der Flüsse - Riesenotter und ihr Lebensraum Regenwald. Knesebeck Verlag München, 1994, 203 Seiten

MacQuarrie, K. (1998). Peru's Amazonian Eden: Manu National Park and Biosphere Reserve. Published by Francis O. Patthey & Sons; 320 pp. Second edition.

MacQuarrie, K. & Bärtschi, A. (2001). Where the Andes meet the Amazon: Peru and Bolivia’s Bahuaja-Sonene and Madidi National Parks. Published by Francis O. Patthey & Sons; 336 pp., ISBN 84-89119-14-7.

Schenck, C. (2000). Among the river wolves. BBC Wildlife Magazine, Nov. 2000; pp. 22-28

Staib E., Schenck C. (1994). Die Familien-Bande. BBC-Wildlife, Nr.12, S. 22-27

Suriname

Duplaix, N. (1978). Synopsis of the status and ecology of the giant otter in Suriname. Otter Trust. Otters 1978; 8 pp.

Duplaix, N. (1980). Giant otters : "Big water dogs in peril". National Geographic Magazine 158(1) 1980; pp. 130-142 PDF 2.5MB

Duplaix, N. (1982). Im Kielwasser der scheuen Riesenotter. Das Tier Tierwelt 2: 4-9

Giant Otters and other animals

Best, R.C. (1984). The aquatic mammals and reptiles of the Amazon. Monographic biol. 56 1984; pp. 371-412

Cabrera, A. & I. Yepes (1940) Mamferos sud-mericanos. Buenos Aires, Comp. Arg. Editores.

Defler. T.R. (1983). Associations of the giant river otter (Pteronura brasiliensis) with freshwater dolphins (Inia geoffrensis). Journal Mammal 64(4) 1983; pp. 692

Emmons, L. (1997). Neotropical Rainforest Mammals. A Field Guide, Second Edition. Illustrations by F. Feer. The University of Chicago Press, Chicago and London

Husson, A.M. (1978). The Mammals of Suriname. Zoologische Monographieen van het Rijksmuseum van Natuurlijke Historie No. 2, published by E.J. Brill, Leiden; 569 pp.

Melquist, W.E. (1984). Status survey of otters (Lutrinae) and spotted cats (Felidae) in Latin America. Report to IUCN College of Forestry, Wildlife and Range Sciences, Univ. of Idaho

Nowak, R.M. (1999) Giant Otter. In Walker's Mammals of the World - 5th Edition Vol. II; pp. 1137-1138

Smith, N.J.H. (1981). Caimans, capybaras, otters, manatees and man in Amazonia. Biol. Conserv. 19; pp. 177-187

de Thoisy, B.,  Catzeflis, F., Richard-Hansen, C.,  Lavergne, A. &  Delcourt, B. (2006). Tapir De Tierras Bajas Y Nutria Gigante : Especies Focales De Los Ecosistemas Amenazados En Guayana Francesa. VII Congresso Internacional Sobre Manejo De Fauna Silvestre Na Amazônia E América Latina. Ilhéus, Bahia, Brasil.

Utreras, V. (2001). Monitoreo Acuático Yasuní - La Nutria Gigante (Pteronura brasiliensis), El Delfín Amazónico (Inia geoffrensis) Y Otras Especies De Mamíferos Acuáticos, En Las Cuencas De Los Ríos Yasuní, Tiputini Y Tivacuno – Parque Nacional Yasuní, Amazonía Ecuatoriana (Período: Enero – Marzo 2001). Informe Técnico (No Publicado). WCS – Programa Ecuador. Quito. 46 Pp.

Utreras, V. (2001). Visión General De Los Mamíferos Acuáticos En El Ecuador, Con Énfasis En El Parque Nacional Yasuní. Pp. 157-164 En: J. P., Jorgenson Y M. Coello Rodríguez (Eds.). Conservación Y Desarrollo Sostenible Del Parque Nacional Yasuní Y Su Área De Influencia. Memorias Del Seminario-Taller 2001. Ministerio Del Ambiente/UNESCO/Wildlife Conservation Society. Editorial Simbioe. Quito, Ecuador.

Utreras, V. (2000). Mamíferos Acuáticos De La Reserva De Producción De Fauna Cuyabeno. Pp. 120-127 En: Yépez, I. (Comp.). Memorias Del Curso Para Guías Naturalistas II De La Reserva De Producción De Fauna Cuyabeno. Ecociencia. Quito.

Neotropical and other otters

Cabello C.C. (1983). La nutria de mar en la Isla de Chiloe. Corporacion National Forestal, Bol. Tec. 6, 1-37.

Davis, J.A. & Duplaix, N. (1976). Neotropical otters. Wildlife, Wld. Conserv. Yb. 1976; pp. 48-52

Estes J. A. (1989). Behavior, ecology and life history of new world otters. Proceedings of the V. International Otter Colloquium. - Habitat 6, Hankensbüttel, 1991.

Gallo J. P. (1989). Distribucion y estado actual de la nutria o perro de agua (Lutra longicaudis annectens Major, 1897) en la Sierra Madre del sur, Mexico.- M.Sc. Thesis. Universidad nacional Autonoma de México, México, D.F

Kruuk, H. (1995). Wild otters - predation and populations. Oxford Univ. Press; 290 pp.

Mason, C.F. & Macdonald, S.M. (1986). Otters - Ecology and Conservation. Cambridge University Press; 236 pp.

Melquist, W.E. (1984). Status survey of otters (Lutrinae) and spotted cats (Felidae) in Latin America. Report to IUCN College of Forestry, Wildlife and Range Sciences, Univ. of Idaho

Parera A. (1993). The neotropical river otter Lutra longicaudis in Iberia Lagoon, Argentina. IUCN Otter Specialist Group Bulletin, 8, pp. 13-16.

Rosas, F. C. W. (2004) Lontra, Lontra longicaudis (Carnivora:Mustelidae). In: Renato Cintra. (Org.). História Natural, ecologia e conservação de algumas espécies de plantas e animais da Amazônia. Manaus: Editora da Universidade Federal do Amazonas, 2004, p. 261-264. PDF 1MB

Utreras, V. And I. Araya. (2002). Distribution And Conservation Status Of The Neotropical Otter (Lutra longicaudis) And Giant Otter (Pteronura Brasiliensis) In Ecuador. Pp. 365-369 In: Dulfer, R., Conroy, J. Nel & A. C. Gutleb (Eds.). Proccedings Viith International Otter Colloquium. Trebon. IUCN Otter Specialist Group Bulletin. Volume 19A, Special Issue.

Utreras, V., M. Rodríguez And I. Araya. (2002). Preliminary Study On The Diet Of The Neotropical Otter (Lutra longicaudis) In The Tiputini River, Yasuni National Park, Ecuadorian Amazonia. Pp. 370-373 In: Dulfer, R., Conroy, J. Nel & A. C. Gutleb (Eds.). Proccedings Viith International Otter Colloquium. Trebon. IUCN Otter Specialist Group Bulletin. Volume 19A, Special Issue

Utreras, V. & D. Tirira. (2001). Nutria Común (Lontra longicaudis). Pp. 57-58 En: D. Tirira (Ed.), Libro Rojo De Los Mamíferos Del Ecuador. SIMBIOE/Ecociencia/Ministerio Del Ambiente/UICN. Serie Libros Rojos Del Ecuador, Tomo 1. Publicación Especial Sobre Los Mamíferos Del Ecuador 4. Quito.

Van Zyll de Jong, C. G. (1972). A systematic review of the Nearctic and Neotropical riverine otters (genus Lutra, Mustelidae, Carnivora). Royal Ontario Museum, Life Sciences, Contribution, 80: 1-104.

TV films and videos

In Search of the Giant Otter (1978, 1984) N. Duplaix, Christopher Parsons, producer. BBC Bristol Natural History Unit 1978, and re-released and re-edited by WGBH-PBS Boston, USA. 50 min.

Im Reich der Riesenotter (1993), A. Hofer, SAT1, 45 min.

Die Riesenotter vom Madre de Dios (1994), C. Schenck, E. Staib, producers N3/ARD, 45 min

The River Wolves (1999), P. Venn, producer. Partridge Films, 45 min

Waika der Otter (? 2000), F.R. de la Fuenta, BR, 45 min

Die Wölfe des Rio Manu – Riesenotter in Peru.( 2001) Felix Heidinger, producer, for ‘Zuflucht Wildnis’ BR/ARD, 45 min

Giant Otters – Wolves of the River (2001). Huw Cordey, producer. BBC ‘Wildlife on One’, 30 mins.

The Miracle of the Pantanal (2004). Satoshi Okabe, Producer. 50-minute film. NHK, Japan.
(Shows many Giant otter cub sequences, including an attack by a Spectacled caiman that fails.)

Websites

www.giantotters.com

Giant Otter Research (this site)
www.giantotterresearch.com

Friends of the Manatee
http://www.amigosdopeixeboi.org.br/
This NGO works with all 5 Amazonian aquatic mammals therefore both Giant Otters and Neotropical Otters are included.

International Otter Survival Fund
http://www.otter.org
This charity works to protect all species of otters worldwide